Photo Credit: Mitchell Coleman

Information for Contributors

Photo by Mitchell Coleman
Photo by Mitchell Coleman

General Information

Updated January 2026

Madroño is a peer-reviewed journal focused on the ecology, systematics, floristics, restoration, and conservation biology of Western American flora, including those of Mexico, Central and South America. Works published in Madroño include (but are not limited to) Research Articles (aka Contents), New Species Descriptions, Taxonomic Revisions, Floristic Works, Regional Checklists, Noteworthy Collections, Notes, and Book Reviews.

Submit manuscripts to Madroño via the PeerTrack website (https://www.editorialmanager.com/madrono/default2.aspx). Contributors submitting a manuscript for the first time must first register online with PeerTrack by following the instructions on the Peer Track website. It is preferred that all authors be members of the California Botanical Society (CBS). Information about the benefits of CBS membership and information on how to join can be found at the CBS website (https://calbotsoc.org).

Manuscripts may be submitted in English or Spanish. English-language manuscripts dealing with taxa or topics of Latin America and Spanish-language manuscripts must have a Spanish RESUMEN and an English ABSTRACT. Manuscripts by authors having outstanding page charges will not be sent for review.

Submitted manuscripts should be accompanied by a cover letter that describes the work, confirms that the submitted manuscript represents the original work of the authors, and describes the extent to which data in the manuscript have been used in other papers that are published, in press, submitted, or soon to be submitted elsewhere. Authors are encouraged to include the names, addresses, and e-mail addresses of two to four potential reviewers with their submitted manuscript.

New Submission Category: Natural History Notes 

Natural history notes are a new submission category beginning with Volume 73 (2026) and will be treated as a ‘Note’ like Noteworthy Collections. We will evaluate the value of these notes based on whether the submission seems like a new, persistent feature or event, rather than a singular event. The note should concern unusual, undescribed, or long-term processes or events of interest to the broader botanical audience. There should be some context so readers can understand the observation and why it is considered unusual or undescribed: why is this considered significant? This type of article should have photographs or figures to accompany the note for clarity.

Submissions should be brief and should include a limited number of photographs, figures, and/or tables that are a core part of the observation. The note should be of scientific interest. Follow author’s guidelines for formal aspects of the manuscript such as citations, and alt-text legends. Include sections on Previous Knowledge and Significance.

New Submission Category: Commentaries

Commentaries should present ideas, concepts, and criticisms that engage readers. That usually means 1) something provocative, 2) something they didn’t know much about, or 3) something they thought they knew something about but found out they were wrong. So, not necessarily a data paper, not necessarily a review paper, but more a “let’s stimulate the audience” paper. These are invited articles, but if you have an idea for a commentary, first contact the editor with a paragraph describing your idea. There is no set structure for this article type but follow our Author Instructions for formal aspects of the manuscript such as citations, and alt-text legends.

 

Author charges for submissions beginning Volume 73

Beginning in 2026, Madroño will participate in a pilot Subscribe to Open (S2O) program through our publishing partner, BioOne. Subscribe to Open is a publishing model that will provide a practical path to opening access to Madroño’s new content while safeguarding the subscription revenue that the California Botanical Society relies upon to fulfill its mission.

What does this mean for our authors and readers? If BioOne’s annual offer is successful, as we expect it to be – meaning that library subscribers continue to support the collection as in the past – then current-year content of Madroño (along with 71 other participating titles) will be made Open Access and freely available to any user worldwide indefinitely.

Because the journal will be online-only and all contributions will be published through the S2O program, we are switching from a system of page charges and optional Open Access fees to standard Article Processing Charges (APCs) as follows: Standard Contents Articles and New Species Descriptions will be $500 for members/$1000 for non-members; Noteworthy Collections and Notes will be $250 for members/$500 for non-members. There is no charge to authors for Book Reviews. All authors must be CalBotSoc members to receive the discounted APCs (or at least three authors if there are more than three authors on the contribution). Author changes to text and figures after typesetting will be charged to authors at $5 per line of text and at cost per figure. There is no additional charge for color figures.

The California Botanical Society is committed to ensuring that all authors have the opportunity to publish in Madroño. To support this goal, we offer reduced article publication charges (APCs) for authors based in certain countries outside the United States, following the discount structure established by Wiley. Eligibility is based on the corresponding author’s affiliation(s) at the time of manuscript submission.

Applicable discounts—whether for members or non-members—will be applied according to the corresponding author’s membership status at time of manuscript acceptance. In addition, student members who are both first and corresponding author at submission receive a 50% reduction in APCs in addition to any country-specific discount.

All discounts are processed automatically by our business office and are independent of the peer-review process. Authors should not communicate with the editor regarding APCs in order to maintain separation between editorial considerations and the author’s ability to pay. Inquiries about APCs can be directed to admin@calbotsoc.org.

Photo by Dean Kelch
Photo by Dean Kelch
Photo by Dean Kelch

Author Instructions

Updated January 2026

Authors are encouraged to follow the format used in recent issues for the type of item submitted (e.g., Research Articles, Noteworthy Collections, etc.). General templates are provided for Research Articles (Contents), Noteworthy Collections, New Species Descriptions, and Book Reviews.

Manuscripts should be submitted as an MS Word file format. Manuscripts must be double spaced throughout, written in 12-point Times New Roman format, and have 1-inch margins. Line numbers are not required; the Peer Track submission system automatically inserts line numbers to facilitate review comments. The order of sections for a Research Article is: Title Page, Abstract (on a its own page), Main text, Acknowledgments, Literature Cited, Tables, Appendices, and Figure Legends (figures are separate image files such as JPG, TIFF, PDF, etc.).

Use of Artificial Intelligence (AI) Policy

Madroño follows the Committee on Publication Ethics (COPE) position that artificial intelligence (AI) tools cannot be recognized as authors, as they cannot take responsibility for the submitted manuscript. Authors are fully responsible for the content of their manuscript, even those parts produced by an AI tool, and are thus liable for any breach of publication ethics. If authors use AI tools such as ChatGPT or other large language models in the writing of a manuscript, in the production of images, or in the collection and analysis of data, they must transparently disclose how and which tool was used in the section of the manuscript to which use of AI pertains, with an additional disclosure statement in the paper’s Acknowledgments. In addition, this must be disclosed in the manuscript cover letter. Tools that are used to improve spelling, grammar, and general editing are excluded from the scope of these guidelines. Authors are responsible for all content generated by AI tools.

Manuscript Content

1.  Title Page.    On title page, provide:

  • The full title (all caps, centered)
  • Authors’ names, affiliations, and mailing addresses. Include the email address for the corresponding author only. Authors with ORCID numbers should add them after their names.
  • Short title to use as a running header— no more than 65 characters (including spaces). This should include author’s last names and an abbreviated title (e.g., Eckert and Sawyer: Foxtail pine importance and conifer diversity).

 

2.  Abstract and Key Words Page

  • On a separate page, provide a one paragraph abstract and up to eight key words or phrases (in alphabetical order). Words from the title may be included in the key words. Each key word should be useful as a term for a literature search.

 

3.  Main Text – Introduction, Methods, Results, and Discussion

  • No heading is needed for the “Introduction”, but this should start on a new page, following the abstract and key words.
  • Major headings should be centered. This includes Methods (or Materials and Methods), Results, Discussion, Acknowledgments (if included), and Literature Cited. Results and Discussion sections may be combined if desired.

 

3.1.  Methods and Results Sections

  • Second-level headings should be centered and in upper and lower case. Example:

Pollination Studies

  • Third-level headings should start the paragraph and be italicized, ending with a period. Example:

            Reproductive isolation. An additional criterion for establishing the taxonomic rank of sympatric taxa …

  • Products used should be cited in parentheses with manufacturer name, city, and state. Examples:

“We used a Li-6800 Portable Photosynthesis System (LiCor Biosciences, Lincoln, NE) to measure…”

“All statistical analyses were conducted in R (R Core Team, R Foundation for Statistical Computing, Vienna, Austria).”

 

3.2.  Presenting the results of statistical analyses

Provide test statistics (e.g., t, F-ratio, etc.), degrees of freedom (if applicable), and P-values for statistical tests [e.g., (P = 0.003) or (P < 0.001)]. If P value is less than 0.0001, just indicate this—it is not necessary to provide all significant figures.

 

3.3.  Other Details

  • Abbreviate figures (Fig. and Figs.) in text and figure captions.
  • Use the whole word “Table” when citing a table in the main text like this “(Table 1)”.
  • Spell out numbers less than 10 (e.g., one, two, three … nine), unless reporting a measurement, range, or aspect of study design (e.g., 7 mm, 2-3 seeds per fruit, Day 5).
  • To use abbreviations, write out the full name at the first instance in follow it with the abbreviation in parentheses. Example: California Native Plant Society (CNPS).
  • Herbarium codes (e.g., JEPS, DAV) may be used without writing out first. See the Index Herbariorum 8th ed. (Regnum Veg. Vol. 120. 1990) for designations of herbarium codes.
  • The following abbreviations may be used without writing out when first introduced: h, min, s, yr, mo, wk, cm, mm, DNA, cpDNA, RNA, dNTP.
  • All measurements and elevations should be in metric units, except specimen citations, which may include English or metric measurements.
  • Designate temperature in degrees Celsius, as in 25°C (use the degree sign, not zero or the letter o).
  • Include a space between all mathematical operators (e.g., +, =)
  • Use the decimal degree representation with positive/negative symbols without degrees symbol, without compass directions (use negative longitude for western North America). For example, the correctly formatted lat/long for UC Berkeley would be 37.871960 -122.259094.

 

4.  Literature Cited

  • Before submitting the manuscript, check each citation in the text against the Literature Cited to see that they match exactly. Delete references that are not cited in the article.
  • All citations must be generally available for access. Unpublished reports or data sets are unacceptable unless they are available as an on-line source. In these instances, the website where the report is available must be included in the reference. Any unpublished information provided to the authors can be cited in the text as a personal communication or observation. In these references: include the source’s affiliation (e.g., L. Smith, Univ. of New Mexico, personal communication). Personal communications are cited in the main text, but not included in the Literature Cited.
  • Check format and capitalization of references before submitting your manuscript. See below for examples.

 

4.1.  In-Text References

Different citations are separated by a semicolon (;) and placed in chronological order. See examples of in-text citations below:

(Johnson 2002; Franklin 2003; Jones 2003)

(Smith 1990; Franklin and Johnson 1995; Jones et al. 2000, 2001)

(Smith in press)

(Jones 1996a, b, c, 1997)

Smith (1990) found that …

 

4.2.  References in the Literature Cited

  • Citations should be in order alphabetically by first author, then grouped by number of authors (1, 2, 3 or more), then alphabetically within each group.
  • For references with many authors, list the first seven authors and then “et al.” If there are eight authors, then list all eight.
  • Type author names in upper and lower case (not all caps). Use formatting examples below.

 

4.2.a  Samples for citation format:

Journal articles:

Critchfield, W. B. 1977. Hybridization of foxtail and bristlecone pines. Madroño 24:193–211.

Rajakaruna, N. and B. A. Bohm. 1999. The edaphic factor and patterns of variation in Lasthenia californica (Asteraceae). American Journal of Botany 86:1576–1596.

Hrusa, F., B. Ertter, A. Sanders, G. Leppig, and E. Dean. 2002. Catalogue of non-native vascular plants occurring spontaneously in California beyond those addressed in The Jepson Manual part I. Madroño 49:61–98.

Journal articles in press:

Jones, R. M. In Press. Studies in the botany of California and parts adjacent. Madroño.

Books:

Wickens, G. E. 2001. Economic botany: principles and practices. Kluwer Academic Publishers, Dordrecht, The Netherlands.

Zar, J. H. 1999. Biostatistical analysis. Prentice Hall, Upper Saddle River, NJ.

Edited Books:

Baker, H. G. and G. L. Stebbins (eds.). 1965. The genetics of colonizing species. Academic Press, New York, NY.

Hickman, J. C. (ed.). 1993. The Jepson manual: higher plants of California. University of California Press, Berkeley, CA.

Book Sections:

Barrett, S. C. H. 2000. Microevolutionary influences of global changes on plant invasions. Pp. 115–140 in H. A. Mooney and R. J. Hobbs (eds.), Invasive species in a changing world. Island Press, Washington, DC.

Sawyer, J. O. and D. Thornburgh. 1988. Montane and subalpine vegetation of the Klamath Mountains. Pp. 699–732 in M. G. Barbour and J. Major (eds.), Terrestrial vegetation of California. California Native Plant Society, Sacramento, CA.

Theses and Dissertations:

Hohn, J. E. 1975. Biosystematic studies of the genus Lewisia, section Cotyledon (Portulacaceae). Unpublished Ph.D. dissertation. University of Washington, Seattle, WA.

Mastrogiuseppe, R. J. 1972. Geographic variation in foxtail pine, Pinus balfouriana Grev. & Balf. Unpublished M.S. thesis. Humboldt State University, Arcata, CA.

Electronic Literature and Databases:

JEPSON FLORA PROJECT (eds.). 2024. The Jepson eFlora. Website https://ucjeps.berkeley.edu/eflora/ [accessed on 5 February 2024].

Johnson, D. R. 1993. Soil survey of Jackson County area, Oregon. United States Department of Agriculture, Soil Conservation Service. Website ftp://ftp-fc.sc.egov.usda.gov/MO1/text_pdf/oregon/or632_text.pdf [accessed 15 May 2023].

Knapp, S. 2011. Rarity, species richness, and the threat of extinction – are plants the same as animals? PLoS Biology 9: e1001067.

Simpson, M. G., K. E. Hasenstab-Lehman and R. B. Kelley. 2021. Johnstonella angelica. In Jepson Flora Project (eds.) Jepson eFlora, Revision 9, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=99828. [accessed on 25 April 2024].

USDA, NRCS. 2004. The PLANTS Database. National Plant Data Center, Baton Rouge, LA. Website http://plants.usda.gov [accessed 04 July 2023].

 

4.3.  Citing iNaturalist Observations

In-Text when referring to 1-10 iNaturalist observations and In-Literature Cited
              For example:
              Lyons, Dean (betweenthelyons). 2022. iNaturalist observation: https://www.inaturalist.org/observations/109511560 [accessed 11 June 2024].
  • If you just want to cite iNaturalist (to refer to it generally, rather than a specific record or set of data), please use the following: iNaturalist. Website: https://www.inaturalist.org. [Accessed DD MONTH YEAR]

 

5.  Tables

  • Tables should be included in manuscript file and place immediately after Literature Cited.
  • Authors must use the MS Word’s table function. Do not use tab and/or hard returns to construct a table. Each table entry should be in its own table cell. Do not use extra returns in table cells. Tables not following these general guidelines will be returned to authors.
  • Tables submitted in other formats will be converted to MS Word.
  • The table legend should be placed immediately above each table. Footnotes are discouraged. Information should be included in the table legend.
  • The first sentence of the table legend should be short and descriptive of the whole table. Additional and/or detailed information follows in subsequent sentences.

Example: Table 1. Short Descriptive Title…

 

6.  Figures and Figure Legends

  • Scales should be included in figures, as should explanation of symbols, including graph coordinates. Maps must include a scale and latitude and longitude or UTM references.
  • Include figure legends in manuscript file and place immediately after the Appendices (or after the tables if there are no appendices)
  • Each figure legend must be complete and informative so that reference to the text is not necessary to understand the content of the figure.
  • The first sentence of the figure legend should be short and descriptive of the whole figure or illustration. Additional and/or detailed information follows in subsequent sentences.

Example: Fig. 1. Short Descriptive Title…

  • For figures with multiple lettered panels, a general title for the figure should be followed by a description of each panel (e.g., “Fig. 5. Fruit structure… A. All fruits. B. Fruits < 0.5 mm.”).

 

7.  Figure and Illustration Files

  • Submit figures and illustrations as separate image files.
  • Acceptable common image formats are GIF, JPEG, TIFF, and PDF. Please check with the editors for other formats.
  • Halftone and color images should be prepared at a minimum of 300 pixels per inch (ppi). Line art illustrations should be prepared at 1200 ppi.
  • Figures should be prepared close to the expected final size in the journal. Maximum page size is 6 inches wide and 8 inches tall. If you are considering a figure that takes the entire length of the page, remember to leave room for a figure caption to fit below all figures.

 

8.  Electronic Supplemental Appendices and Archived Data

Authors have the opportunity to publish online appendices with papers through BioOne. References to supplemental data sets archived elsewhere are designated with an “S” (e.g., Appendix S1). Print appendices may also be published directly in the printed (and PDF) form of the journal. References to printed appendices are simply numbered (e.g., Appendix 1).

If the authors are archiving their data in an online repository, such as Dryad, a “Data Accessibility” statement should be inserted after the Acknowledgments and before the Literature Cited. The statement should provide the name of the database, digital object identifiers, and stable URLs.

 

9.  Use of Common and Taxonomic Names

  • Common names are written out with the first letter capitalized, including words that are not proper nouns. Examples: Coast Redwood, Monterey Pine.
  • Identify organisms by full botanical name at first use (including the taxonomic authority and family name) in the text and in the abstract. This may be followed by common name in parentheses, if desired. Authorities for botanical names are not required for the title. The Latin name of the taxon is italicized. Taxonomic authorities must be confirmed in the Jepson eFlora or Tropicos.org.
  • Genus names are spelled out the first time they are used and should be abbreviated thereafter (unless starting a sentence or where abbreviations may be confusing). If an article contains multiple botanical names, spell out the genus name the first time it appears in each major section.
  • Avoid or minimize using the name of a new taxon until after it appears at the beginning of the formal description. It is especially important to avoid using the names of new taxa in the title and short title of the manuscript.

 

10.  Accessibility Guidelines: Providing Alt Text for Images 

To ensure your work is accessible to the widest possible audience, including readers who use screen readers or other assistive technologies, Madroño requires all authors to provide alternative text (alt text) for all figures, tables, images and visual elements in your manuscript.

What is Alt Text?

Alt text is a brief description of an image that is embedded in the digital content and read aloud by screen readers. It helps readers with visual impairments understand the content and purpose of images that they cannot see. It is a required component to all Madroño submissions after June 28, 2025.

Why is Alt Text Important?

Providing alt text is crucial for accessibility. It ensures that all readers can access the important information conveyed by images, enabling them to fully comprehend and engage with your work. Additionally, meeting accessibility standards is becoming increasingly important for publication and distribution.

How to Write Effective Alt Text:

  • Keep it Concise: Alt text should be short and to the point. Aim for a concise description that conveys the essence of the image’s content and purpose. Some suggest aiming for around 50 words or less, while avoiding descriptions that are overly lengthy or burdensome for screen reader users.

  • Focus on Relevant Information: Describe the image in the context of your manuscript. Consider what information is essential for a reader who cannot see the image to understand its significance in the text. Alt text is not the figure caption, nor table title.

  • Be Objective: Stick to describing what is visually present in the image and avoid subjective interpretations or emotional language.

  • Avoid Redundancy: Do not repeat information already provided in the figure caption, table title or associated text. Screen readers will read the figure captions, table titles and the alt text, so focus on supplementing rather than duplicating existing information. What do you want the reader to take away from this figure or table if they can’t see it.

  • Format Correctly: Do not use formatting like bullet points or lists within the alt text. Conclude the alt text with a period to allow for a pause in the screen reader.

  • Spell Out Abbreviations and Symbols: Avoid acronyms and symbols that screen readers might misinterpret. Spell out the full words for clarity.

  • For Decorative Images: If an image is purely decorative and does not convey any important information, you can indicate this by using “decorative” in the alt text (e.g., “decorative, no alt text needed”).

Submitting Alt Text:

Please place Alt-text directly below each regular caption, preceded by “[Alt-Text].” For Tables provide alt-text as [Table Summary] and give a brief summary of the table. This is required during manuscript submission.

Here are examples:
Figure 1 Measurement device and the AA5083 specimen Source: Author’s own work

[Alt-Text] The image shows a periodic push and pull electric device and the comparative distribution of seawater level.

For Table Alt-text, please use wording [Table Summary]. Example:

Table 1. Seed bank densities of Arctostaphylos species.

[Table Summary] The densities of ten Arctostaphylos soil seed banks are listed. Also included are fruit diameters, fruit volumes, and the number of seeds in two soil layers. The numbers are averages with standard errors of the mean in parentheses. The ratio L/U refers to the lower 2-5 cm layer (L) divided by the upper 0-2 cm layer (U).

Table 2. Coefficients for the top GLMM model for each genus.

[Table Summary] Phenological GLMM model coefficients for two genera are listed and whether they were significant or not in the model.

Thoughtful and accurate alt text contributes to a more inclusive and accessible reading experience. For questions about writing alt text, contact the Madroño Editor at madronoeditor@gmail.com or Managing Editor managingeditormadrono@gmail.com.

Photo by Dean Kelch
Photo by Dean Kelch
Photo by Dean Kelch
Photo by Dean Kelch
Photo by Dean Kelch
Photo by Dean Kelch
Photo by Dean Kelch
Photo by Mitchell Coleman
Photo by Mitchell Coleman
Photo by Mitchell Coleman
Photo by Mitchell Coleman
Photo by Mitchell Coleman
Photo by Mitchell Coleman
Photo by Mitchell Coleman
Photo by Mitchell Coleman
Photo by Mitchell Coleman
DCIM100MEDIADJI_0115.JPG
Photo by Mitchell Coleman
Photo by Mitchell Coleman
Photo by Mitchell Coleman

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c/o Jepson Herbarium 1001 Valley Life Sciences Building Berkeley, California 94720-2465

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